IAL Biology • Topic 2 Review

Cell Division, Meiosis & Reproduction

Chapter Review — Cell Cycle, Mitosis, Meiosis, Gametes & Fertilisation

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Diagnostic Quiz

Test what you already remember before we begin. Choose the best answer for each question.

1. What is the correct order of the cell cycle?
The cell cycle proceeds: Interphase (G1, S, G2) → Mitosis → Cytokinesis.
2. How many daughter cells does meiosis produce?
Meiosis produces 4 genetically different haploid daughter cells.
3. During which phase of interphase is DNA replicated?
DNA replication occurs during the S (synthesis) phase of interphase.
4. What is a gamete?
A gamete is a haploid sex cell (sperm or egg in animals; pollen or egg cell in plants).
5. In flowering plants, what does double fertilisation produce?
In double fertilisation, one male nucleus fuses with the egg to form a zygote, and the other fuses with two polar nuclei to form the triploid endosperm nucleus.
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Learning Objectives

Specification Checklist

Click each objective as you feel confident with it.

  • Describe the cell cycle as a regulated process of interphase, mitosis and cytokinesis, producing two identical daughter cells
  • Explain the replication and separation of genetic material in the main stages of mitosis
  • Explain the importance of mitosis in growth, repair of damaged tissues and asexual reproduction
  • Describe the role of meiosis in the production of haploid gametes, including its stages
  • Explain the replication and separation of genetic material in the main stages of meiosis
  • Explain how meiosis results in genetic variation through independent assortment and crossing over
  • Describe the development of female and male gametes in mammals and plants (pollen grain and embryo sac)
  • Describe the adaptations of gametes for their functions
  • Describe fertilisation in mammals and double fertilisation in plants, including the roles of the tube nucleus and generative nucleus
  • Explain how random fertilisation during sexual reproduction brings about genetic variation
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Tier 1 | Target: C Grade | AO1: Knowledge & Recall

Foundation Recall

Flashcard Retrieval Practice

Tap each card to reveal the definition. Try to recall the answer before flipping.

Cell Cycle
The regulated sequence of events in which a cell grows, replicates its DNA and divides. It consists of interphase, mitosis and cytokinesis.
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Interphase
The longest phase of the cell cycle, subdivided into G1 (cell growth), S phase (DNA replication) and G2 (preparation for division). The cell is metabolically active.
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Mitosis
Nuclear division producing two genetically identical daughter nuclei, each with the same number of chromosomes as the parent cell (diploid). Stages: prophase, metaphase, anaphase, telophase.
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Cytokinesis
The division of the cytoplasm following nuclear division, producing two separate daughter cells. In animal cells the membrane pinches inward; in plant cells a cell plate forms.
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Meiosis
A type of nuclear division that produces four genetically different haploid daughter cells from a single diploid cell. It involves two successive divisions: meiosis I and meiosis II.
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Haploid (n)
A cell or nucleus containing one complete set of chromosomes — half the diploid number. Gametes are haploid. In humans, n = 23.
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Diploid (2n)
A cell or nucleus containing two complete sets of chromosomes, one from each parent. Body (somatic) cells are diploid. In humans, 2n = 46.
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Gamete
A haploid reproductive cell (sex cell) that fuses with another gamete during fertilisation to form a diploid zygote. Examples: sperm, egg, pollen grain.
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Homologous Chromosomes
A pair of chromosomes of the same size, with the same centromere position and the same genes at the same loci, but possibly different alleles. One is inherited from each parent.
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Crossing Over
The exchange of genetic material between non-sister chromatids of homologous chromosomes during prophase I of meiosis. Produces recombinant chromosomes, increasing genetic variation.
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Independent Assortment
The random orientation of homologous pairs at the metaphase plate during meiosis I. This means each gamete receives a random combination of maternal and paternal chromosomes.
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Fertilisation
The fusion of two haploid gamete nuclei to form a diploid zygote. This restores the diploid chromosome number and introduces genetic variation.
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Pollen Grain
The male gametophyte in flowering plants. Contains a tube nucleus (controls pollen tube growth) and a generative nucleus (divides to form two male gamete nuclei).
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Embryo Sac
The mature female gametophyte in flowering plants. Contains the egg cell, two synergid cells, three antipodal cells and a large central cell with two polar nuclei.
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Zygote
A diploid cell formed by the fusion of two haploid gametes during fertilisation. It is the first cell of a new organism.
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Fill in the Blanks

Use the word bank to complete the passage below.

interphase mitosis cytokinesis S phase haploid meiosis four crossing over independent assortment diploid homologous gametes

The cell cycle consists of three main stages. The first and longest stage is , during which DNA is replicated in the . The nucleus then divides by , and the cytoplasm splits during . This produces two genetically identical daughter cells.

To produce sex cells called , cells divide by . This produces genetically different cells. In prophase I, chromosomes pair up and exchange genetic material in a process called . The random alignment of chromosome pairs during metaphase I is known as .

Key Term Matching

Click a term on the left, then click its matching definition on the right.

Terms
Prophase
Metaphase
Anaphase
Telophase
Chiasma
Bivalent
Definitions
Centromeres split and chromatids are pulled to opposite poles
A pair of homologous chromosomes joined together during meiosis I
Chromosomes condense, nuclear envelope breaks down, spindle forms
The point where non-sister chromatids exchange material during crossing over
Chromosomes reach the poles, nuclear envelopes reform, chromosomes decondense
Chromosomes line up on the equator of the cell, attached to spindle fibres

Tier 1 Quick Quiz

1. State the function of cytokinesis.
Cytokinesis is the division of the cytoplasm to form two separate daughter cells after nuclear division.
2. How many chromosomes does a human gamete contain?
Human gametes are haploid (n) and contain 23 chromosomes.
3. Which type of cell division is used for growth and repair?
Mitosis produces genetically identical cells, essential for growth and repair of tissues.
4. What is the name of the structure formed when homologous chromosomes pair up in meiosis I?
When homologous chromosomes pair up (synapsis) during prophase I, the structure is called a bivalent (or tetrad).
5. In a pollen grain, what is the role of the generative nucleus?
The generative nucleus divides by mitosis to produce two male gamete nuclei, which are involved in double fertilisation.
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Tier 2 | Target: C–B Grade | AO2: Application & Understanding

Application & Understanding

Interactive Cell Cycle Diagram

Click on each phase of the cell cycle to reveal a description.

G1 S G2 M C CELL CYCLE INTERPHASE

Click a phase in the diagram above to learn about it.

Mitosis vs Meiosis Comparison

Click on each cell in the table to reveal the answer. Test yourself first!

FeatureMitosisMeiosis
Number of divisionsClick to revealClick to reveal
Number of daughter cellsClick to revealClick to reveal
Chromosome number of daughtersClick to revealClick to reveal
Genetic variation?Click to revealClick to reveal
Where it occursClick to revealClick to reveal
PurposeClick to revealClick to reveal
Homologous pairing?Click to revealClick to reveal
Crossing over?Click to revealClick to reveal

Sequence the Stages of Meiosis

Click the stages in the correct order (1st to 8th). Click “Reset” to start again.

Prophase II — chromosomes condense again, no crossing over
Metaphase I — bivalents line up randomly on the equator
Anaphase II — centromeres split, sister chromatids pulled apart
Telophase I — nuclear envelopes reform, cell divides (cytokinesis I)
Prophase I — chromosomes condense, bivalents form, crossing over occurs
Telophase II — four haploid daughter cells formed
Anaphase I — homologous chromosomes pulled to opposite poles
Metaphase II — chromosomes line up on equator of each cell

Explain Questions (3–4 marks)

Q1. Explain why mitosis is important for the growth of a multicellular organism. (3 marks)

Use these sentence starters to structure your answer:

Mitosis is important for growth because…
The daughter cells produced are… which means…
This ensures that all new cells have…

Mitosis produces two genetically identical daughter cells with the same diploid number of chromosomes as the parent cell. This is important for growth because the organism needs to increase its cell number while ensuring every new cell contains the complete set of genetic instructions needed to function correctly. Because the daughter cells are genetically identical, the same proteins can be produced, allowing the new cells to carry out the same functions as the original cell.

Q2. Explain how crossing over during meiosis leads to genetic variation. (3 marks)

During prophase I, homologous chromosomes…
Non-sister chromatids exchange… at points called…
This results in chromatids with… which means the gametes…

During prophase I of meiosis, homologous chromosomes pair up to form bivalents. Non-sister chromatids wrap around each other and exchange sections of DNA at points called chiasmata. This means the chromatids now have new combinations of alleles that were not present in either parent chromosome. The resulting gametes are therefore genetically different from each other and from the parent cell, increasing genetic variation in the offspring.

Application Scenarios

Scenario: A gardener takes a cutting from a plant and grows it into a new individual. What type of cell division is responsible for the growth of this new plant, and why is the offspring genetically identical to the parent?

The cutting grows by… because new cells are needed for…
The offspring is genetically identical because mitosis produces…

The cutting grows into a new plant by mitosis, because new cells are needed for growth and development. Mitosis produces daughter cells that are genetically identical to the parent cell, as the DNA is replicated exactly during the S phase and then separated equally during nuclear division. Since only one parent is involved and no gametes or fertilisation occur, this is asexual reproduction, and the offspring is a genetic clone of the parent plant.

Scenario: After a skin wound, new cells are produced to repair the damaged tissue. Explain why mitosis, not meiosis, is used for this repair.

Mitosis is used because the replacement cells must be genetically identical to the original skin cells so they can carry out the same functions (e.g. producing the same structural proteins such as keratin). Mitosis produces diploid cells with the full set of 46 chromosomes. Meiosis would produce haploid cells with only 23 chromosomes, which would not have all the genes necessary to function as skin cells. Additionally, meiosis introduces genetic variation, which is not desirable when replacing specific tissue cells.

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Tier 3 | Target: B–A Grade | AO3: Analysis & Evaluation

Analysis & Evaluation

Extended Response Practice (6 marks)

Evaluate the importance of meiosis in maintaining genetic variation within a population. (6 marks)

Plan your answer using this checklist:

  • Define meiosis and state what it produces
  • Explain crossing over and its effect on allele combinations
  • Explain independent assortment and its effect
  • Link to random fertilisation as a further source of variation
  • Evaluate — why is genetic variation important for survival/natural selection?
  • Conclude with a judgement about the overall significance of meiosis

Meiosis is a form of nuclear division that produces four genetically different haploid gametes from a single diploid parent cell. It is the primary source of genetic variation in sexually reproducing organisms.

During prophase I, homologous chromosomes pair up to form bivalents and crossing over occurs. Non-sister chromatids exchange sections of DNA at chiasmata, producing recombinant chromatids with new combinations of alleles. This means each gamete can carry a unique set of alleles that was not present in the parent.

During metaphase I, homologous pairs align randomly on the metaphase plate in a process called independent assortment. Because each pair can orient in two ways, a human cell (with 23 pairs) can produce 223 (over 8 million) different combinations of maternal and paternal chromosomes in the gametes.

When combined with random fertilisation — any sperm can fuse with any egg — the total number of possible genetic combinations becomes enormous (223 × 223 = over 70 trillion), and this is before accounting for crossing over.

This genetic variation is critically important for the survival of a population. It provides the raw material for natural selection: in a changing environment, some individuals will have allele combinations that confer a selective advantage, enabling them to survive and reproduce. Without meiosis, offspring would be genetically identical to their parents, and a population could be wiped out by a single disease or environmental change.

In conclusion, meiosis is fundamentally important for maintaining genetic diversity. Through crossing over, independent assortment and its link to random fertilisation, it ensures that each generation contains genetically unique individuals, enabling populations to adapt and evolve over time.

Evaluation Task

Some organisms reproduce only asexually (by mitosis). Evaluate whether this is an advantage or disadvantage compared to sexual reproduction involving meiosis. (6 marks)

Advantages of asexual reproduction: It is faster as no mate is needed; all individuals can reproduce; offspring are genetically identical to the parent, which is advantageous in stable environments where the parent is already well-adapted; less energy is expended as there is no need to produce gametes or find a mate.

Disadvantages of asexual reproduction: No genetic variation is produced (no crossing over, independent assortment or random fertilisation). This means the population is vulnerable to environmental change — a single disease or change in conditions could eliminate the entire population. There is no raw material for natural selection to act upon, so the population cannot evolve and adapt over time.

Evaluation: Overall, while asexual reproduction is advantageous in stable environments for rapid population growth, sexual reproduction (involving meiosis) provides a significant long-term advantage because the genetic variation produced allows populations to adapt to changing conditions and resist pathogens. This is why the vast majority of complex organisms use sexual reproduction.

Data Analysis

The table below shows chromosome numbers for different organisms. Use the data to answer the questions.

OrganismDiploid number (2n)Haploid number (n)
Human4623
Fruit fly84
Potato4824
Dog7839
Wheat4221

Questions:

(a) Calculate the number of possible chromosome combinations in the gametes of a fruit fly due to independent assortment alone. Show your working.

Number of pairs (n) = 4. Possible combinations = 2n = 24 = 16 different combinations.

(b) A dog has a diploid number of 78. If two dogs mate, calculate the total number of possible zygote combinations due to independent assortment alone (ignoring crossing over).

n = 39. Each parent can produce 239 different gametes. Total combinations = 239 × 239 = 278 ≈ 3.0 × 1023 possible combinations. This is an astronomically large number, illustrating the enormous genetic diversity possible from independent assortment and random fertilisation alone.

(c) Explain why the actual number of genetically different gametes produced by an organism is even greater than the value calculated from independent assortment alone.

The calculation 2n only accounts for independent assortment — the random orientation of homologous pairs. Crossing over during prophase I creates additional variation by exchanging sections of DNA between non-sister chromatids, producing recombinant chromosomes with new allele combinations. Since crossing over can occur at different points along the chromosomes and involves different chromatids, the number of genetically different gametes is far greater than 2n.

Synoptic Links

Think Deeper: Linking Cell Division to DNA Replication

Links to: Ch 2B — DNA Replication Links to: Ch 2C — Mutations

Question: DNA replication occurs during the S phase of interphase before both mitosis and meiosis. Explain why accurate DNA replication is essential before mitosis, and discuss what could happen if errors (mutations) occur during this process.

During the S phase, DNA replication occurs by the semi-conservative mechanism: the double helix unwinds, hydrogen bonds between complementary base pairs break, and each strand acts as a template. DNA polymerase adds free nucleotides by complementary base pairing (A-T, C-G), producing two identical copies of each DNA molecule.

Accurate replication is essential before mitosis because the purpose of mitosis is to produce genetically identical daughter cells. If replication errors occur and are not corrected by proofreading enzymes, mutations may be introduced. These mutations will be passed to both daughter cells and all subsequent cells produced from them.

If a mutation affects a gene controlling the cell cycle (e.g. a proto-oncogene or tumour suppressor gene), the cell may divide uncontrollably, leading to tumour formation and potentially cancer. This illustrates how the accuracy of DNA replication is fundamentally linked to the proper regulation of cell division.

Think Deeper: Cell Division and Cell Ultrastructure

Links to: Ch 3A — Cell Ultrastructure

Question: Explain the role of the centrioles and the spindle fibres (microtubules) during mitosis. Why are centrioles absent in most plant cells, and how do plant cells still manage to divide?

In animal cells, centrioles move to opposite poles during prophase and organise the spindle fibres (composed of microtubules made of the protein tubulin). Spindle fibres attach to centromeres of chromosomes at the kinetochore and are responsible for moving chromosomes during anaphase.

Most plant cells lack centrioles but can still form a spindle. They use microtubule organising centres (MTOCs) at each pole to nucleate spindle fibres. The process of chromosome separation is functionally the same. Additionally, during cytokinesis, plant cells cannot pinch inward due to their rigid cell wall; instead, vesicles from the Golgi apparatus fuse at the equator to form a cell plate, which develops into a new cell wall.

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Exam Practice

2-Mark Question

State two differences between mitosis and meiosis. (2 marks)

Mitosis produces 2 daughter cells; meiosis produces 4 daughter cells (1 mark)
Mitosis produces genetically identical / diploid cells; meiosis produces genetically different / haploid cells (1 mark)

Accept: crossing over occurs in meiosis but not mitosis; homologous pairing occurs in meiosis but not mitosis; mitosis involves one division, meiosis involves two.

4-Mark Question

Describe the process of double fertilisation in flowering plants. (4 marks)

Pollen grain lands on stigma; pollen tube grows down the style towards the ovule, controlled by the tube nucleus (1 mark)
The generative nucleus divides by mitosis to produce two male gamete nuclei (1 mark)
One male gamete nucleus fuses with the egg cell nucleus to form a diploid zygote (which develops into the embryo) (1 mark)
The other male gamete nucleus fuses with the two polar nuclei in the central cell to form the triploid (3n) endosperm nucleus (which develops into a food store) (1 mark)

6-Mark Question

Describe and explain how the structure of a mammalian sperm cell and egg cell are adapted for their roles in fertilisation. (6 marks)

Plan your answer:

  • Sperm: streamlined shape, acrosome, many mitochondria, flagellum/tail, haploid nucleus
  • Egg: large size, cytoplasm with nutrients, cortical granules, haploid nucleus, zona pellucida
  • Link each structural feature to its function

Sperm cell adaptations:

The sperm cell has a streamlined shape to reduce friction as it swims through the female reproductive tract. It possesses a long flagellum (tail) which enables it to swim towards the egg. The mid-piece contains many mitochondria arranged in a spiral, which carry out aerobic respiration to release the ATP needed to power the movement of the flagellum. The acrosome at the head contains hydrolytic enzymes (e.g. hyaluronidase and acrosin) that digest the protective layers (zona pellucida and follicle cells) surrounding the egg, allowing the sperm to penetrate. The nucleus is haploid (n) so that when it fuses with the haploid egg, the diploid number (2n) is restored.

Egg cell adaptations:

The egg cell is much larger than the sperm because it contains a large volume of cytoplasm rich in nutrients (lipid droplets, proteins) to nourish the early embryo after fertilisation. It has a haploid nucleus to ensure the correct diploid number upon fertilisation. The egg is surrounded by a zona pellucida (glycoprotein layer) which contains receptors for sperm binding and prevents polyspermy. After fertilisation, cortical granules release their contents, causing the zona pellucida to harden (the cortical reaction), blocking further sperm entry and ensuring only one sperm fertilises the egg.

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Final Assessment

Test your understanding across all three assessment objectives. These questions are more challenging than the diagnostic.

1. A cell in G2 of interphase has 24 chromosomes. How many DNA molecules does it contain?
After the S phase, DNA has been replicated. Each chromosome consists of two sister chromatids joined at the centromere, so 24 chromosomes × 2 = 48 DNA molecules.
2. Which of the following events occurs in meiosis but NOT in mitosis?
Bivalent formation (synapsis) and crossing over are unique to meiosis I. The other events occur in both mitosis and meiosis.
3. In a species with a diploid number of 6, how many different gamete combinations are possible from independent assortment alone?
2n = 6, so n = 3. Possible combinations = 2n = 23 = 8.
4. During double fertilisation in plants, the endosperm nucleus is:
One male gamete nucleus (n) fuses with two polar nuclei (n + n) = triploid endosperm nucleus (3n).
5. A farmer breeds genetically identical strawberry plants using runners (a form of asexual reproduction). Which statement best evaluates the risk of this approach?
Without genetic variation, all individuals have the same susceptibility. This is a key evaluation point linking asexual reproduction, mitosis and genetic vulnerability.
6. Which of the following correctly describes the roles of the tube nucleus and generative nucleus in a pollen grain?
The tube nucleus directs the growth of the pollen tube down the style. The generative nucleus undergoes mitosis to produce two male gamete nuclei for double fertilisation.
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Confidence Check

How confident do you feel about this topic now? Select one.

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Vocabulary Grid

Cell Cycle
The regulated sequence of interphase, mitosis and cytokinesis in which a cell grows and divides.
Interphase
The phase of the cell cycle where the cell grows and replicates its DNA (G1, S, G2 sub-phases).
G1 Phase
First growth phase of interphase; cell grows, organelles replicate, proteins are synthesised.
S Phase
Synthesis phase; DNA replication occurs, each chromosome becomes two sister chromatids.
G2 Phase
Second growth phase; cell checks replicated DNA and prepares for mitosis (e.g. synthesises spindle proteins).
Mitosis
Nuclear division producing two genetically identical diploid nuclei. Stages: PMAT.
Cytokinesis
Division of the cytoplasm to form two separate daughter cells after nuclear division.
Prophase
Chromosomes condense and become visible; nuclear envelope breaks down; spindle fibres form.
Metaphase
Chromosomes align on the cell equator (metaphase plate); spindle fibres attach to centromeres.
Anaphase
Centromeres split; sister chromatids are pulled to opposite poles by spindle fibres.
Telophase
Chromosomes reach the poles and decondense; nuclear envelopes reform around each set.
Meiosis
Nuclear division producing four genetically different haploid cells from one diploid cell; two divisions (I and II).
Haploid (n)
Having one complete set of chromosomes. Gametes are haploid.
Diploid (2n)
Having two complete sets of chromosomes, one from each parent. Body cells are diploid.
Homologous Chromosomes
A matching pair of chromosomes (same size, same gene loci) with one from each parent.
Bivalent
A pair of homologous chromosomes associated together during prophase I of meiosis.
Crossing Over
Exchange of DNA between non-sister chromatids of homologous chromosomes during prophase I.
Chiasma (pl. chiasmata)
The point where crossing over occurs between non-sister chromatids.
Independent Assortment
Random orientation of homologous pairs during metaphase I, leading to random combinations in gametes.
Gamete
A haploid sex cell that fuses with another gamete at fertilisation (e.g. sperm, egg, pollen).
Zygote
A diploid cell formed by fusion of two haploid gametes; the first cell of a new organism.
Fertilisation
Fusion of two haploid gamete nuclei to form a diploid zygote, restoring the diploid number.
Pollen Grain
Male gametophyte in plants; contains tube nucleus and generative nucleus.
Tube Nucleus
Controls the growth of the pollen tube down through the style towards the ovule.
Generative Nucleus
Divides by mitosis to form two male gamete nuclei for double fertilisation.
Embryo Sac
Mature female gametophyte; contains the egg cell, polar nuclei, synergids and antipodal cells.
Double Fertilisation
In plants: one male nucleus fuses with egg (zygote), the other fuses with polar nuclei (triploid endosperm).
Endosperm
Triploid (3n) nutritive tissue formed from fusion of male gamete nucleus with two polar nuclei.
Acrosome
Structure at the head of a sperm cell containing enzymes to digest the egg's protective layers.
Cortical Reaction
Release of cortical granule contents after fertilisation, hardening the zona pellucida to prevent polyspermy.